biologe

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Tag: Acari

Oribatida mites: Fast runners and slow crawlers

Microhabitats often consist of a complexity of organism species. Under suitable conditions, samples can be kept „alive“ for months and even for years by regularly adding moisture and organic tissue, in case of my sample of this footage: patato pieces.

 

 

Mites of the Oribatida and their different ways of locomotion. Copyrights: Stefan F. Wirth, Berlin April 2019. Please give the video a like on youtube too.

 

Soil samples from island Norderney

 

This soil sample was collected in summer 2018 on the North Sea island Usedom during my participation at the „Geo Tag der Natur“. It contained several specimens of the predatory chilopode Lithobius sp. and pieces of rotting wood, moss and forestground, everything collected under rotting treetrunks and tree branches. The samples additionally contained the carabid beetle Pterosticus cf. niger and ants of genus Lasius. Samples were collected in a small forest area with wetland aspects. The soil quality was rather moist.

 

Astigmatid mites

 

I later added potato pieces and regularly some water droplets to the sample with still living big arthropods/ insects. After some weeks, specimens of the astigmatid mite Acodyledon cf. schmitzi developed on dryer areas of the potato pieces. These mites were presumably phoretic associates of the carabid beetles. They died out after several months, after the sample had dried out a little bit and may be due to changes of the room temperature during winter time.

 

Oribatida

 

Now, almost a year later, the micro habitat is inhabited by mites of the Oribatida in greater numbers of specimens of at least three species: Nothrus sp. (genus not yet clarified), Nothrus palustris (already found for the first time shortly after the sample collection) and a species of Phthiracarida.

 

Locomotion and biodiversity

 

Purpose of the short film is to show different organisms, cultured after about a year in this sample: mites, nematodes, collembolans and microorganisms, fungae and bacteria. Of the bigger arthropods/insects, only one Lithobius species survived until now.  Also the diversity of ways of locomotion in different oribatid species is emphasized: There are slow crawlers (Nothrus) and fast runners (Phthiracarida).

 

Berlin, April 2019, Copyrights Stefan F. Wirth

Microscopic wrack inhabitants: Mites (Ameronothridae), Protozoans, nematodes and Dipterans

Decomposing detritus (mostly dead algae debris) of marine organic material, laying onshore more or less close to the water line, containing seaweed or cadavers of aqatic animals, is named wrack. Wrack can appear under different kinds of ecological circumstances. In case, it would be in permanent contact with sea water, it might be mostly decomposed by marine organisms. But due to different reasons, wrack can land apart from a permanent sea water contact or even no sea water contact at all any more.

Here mostly terrestrial organisms with a tolerance for salty conditions would inhabit and decompose this piece of detritus. Sandhoppers (Cristacea) are known to switch between wracks of different conditions. They can for example carry mites or nematodes from one wrack habitat to another. Dead organic material generally always needs to be decomposed by living organisms, otherwise the whole ecological system would be harmed.

 

A specific kind of micro habitats

 

A small habitat, which would dry out after a while and thus exists only for a limited time, is called ephemere biochorion. Organisms being adapted to live there, must have adaptations, to leave their habitat by time to avoid desiccation. One option is a life strategy, which is named phoresy. Weaker organisms, unable to desperse themselves efficiently use other animals, such as winged insects, to take a ride on them to new habitats with suitable conditions for a development. Generally phoretic organisms can for example be represented by different groups of mites (e.g. Uropodida, Gamasina, Tarsenomidae, Scutacaridae, some Oribatida, Astigmata) and nematodes (Rhabditida).

 

Mites and nematodes

 

In case of wrack, decomposing close to the waterline, but without or only occasional water contact, Pellioditis marina (Nematoda, Rhabditida) is for example known as phoretic inhabitant along German coasts. Worldwide, crypitical sibling species of P. marina were meanwhile discovered. Depending on the exact situation of the wrack, also aquatic nematodes could appear there for a while. I couldn’t determine the nematode in my footage unfortunately at all, because I did not prepare slides of them enable a larger microscopic magnification. Phoretic mites can be associated with sand-hoppers (Amphipoda, Crustacea) and thus appear in wrack. Mites of the Histiostomatidae (Astigmata) were for example discovered in such a context by some researchers.

 

Mites of the Ameronothridae (Oribatida), sand-hoppers and dipterans

 

I so far never found them randomly, but also didn’t explicitely seek for histiostomatid species until now. My sample did not contain any Astigmata or I at least didn’t find them. Common inhabitants of decomposing wrack are oribatid mites of the Ameronothridae. This taxon with a worldwide distribution is charaterized by specific adaptations to deal as terrestrial organisms with (partly extreme) salty marine conditions. They are mostly algae feeders. Some species are known to appear in wrack. The sample, which I collected in context of the so called „Geo Tag der Natur 2018“ (Geo (journal) day of nature) in Norddeich Mole (East Frisian coast of Germany) contained many specimens (ca. 40, sample size of about 20×20 cm) of the Ameronothridae-species Ameronothrus sp.. My footage shows only one living specimen, as all had died until I began my filming activities.

Inhabitants of decomposing algae tissue along a beach at German North Sea, all copyrights Stefan F. Wirth

 

But I preserved several dead specimens for scientifc purposes. Ameronothridae might, according to literature, use phoresy via birds, but also might disperse themselves over smaller distances, due to their well developed cuticle, protecting against desiccation, and their rather fast locomotion abilities. Larvae of different species of flies (Diptera) developed inside my sample and hatched under my laboratory conditions after about two weeks. They intensively contributed to a fast decomposition of that organic marine tissue. Sand-hoppers were by the way not found at all.

 

Bacteria and protozoans

 

Bacteria are most important decomposers. But the function of protozoans (here e.g. Ciliata) in regard to the process of wrack degradation, which could still be isolated alive after about two weeks of decomposition,  is unknown to me. My sample was found almost on top of a dike, meters away from the highest tide in that area and consisted mostly of the seaweed Fucus vesiculosus.It also contained sea gull feathers.

 

Berlin/ Norddeich Mole June/August/November 2018 Copyrights Stefan F. Wirth

The mite Histiostoma blomquisti and creationism in Louisiana/ USA

New Orleans is a dynamic and a very lively city, a city full of freedom and tolerance, a city of life style and of a unique cuisine. A mix-up of ethic groups and cultures, New Orleans, the modern city, a world metropolis.

But this is not Louisiana, it is an island, an exception, New Orleans is not even the capital of this Southern Federal State. The rest of Louisiana is landscape, swamps, wetlands, pine woods, red sand, even a Red River, harmless snakes and a touch of music, not any kind of music, Louisiana is the birthplace of Jazz.

LA is unfortunately also home land of a strict two class society, with the white race in a top position and the native Americans and the blacks on a level much further below. The latter inhabit usually the so called „no go-areas“. That’s where poverty lives, where a permanent existential emergency dominates the daily routine, and yes, where based on all this distress also criminality finds a new home again and again.

It’s a land, where racism is still alive, where colored people take over the minor jobs, while the whites reign over in high positions. A land of injustice, of inconsistency, a land of religious fanaticism.

Nevertheless, beauty can be found everywhere. When the setting sun illuminates the colorful water of the Red river or shines on lying around rusty metal scrap, then a  spectacle of glowing colors blinds the eyes of the audience. When I walked across the fields and forests, then I found an inspiring silence, a flood of harmonic nature impressions, giant millipedes of Polydesmida hiding under freely lying stone chunks, butterflies colonizing rotting fruits and fluttering with a gentle noise in the air, colorful water turtles taking their sun baths around ponds, and under suitable weather conditions I witnessed wedding celebrations of a very special kind: winged ants rose in the air to mate for their first and only time.

The most common ant species is the Red Imported fire Ant, Solenopsis invicta, a fascinating social insect, but far away from being native to Louisiana. It is a so called invasive species, which was transferred to the Southern US via ballast substrate of ships coming from Southern America. The high frequency of ant colonies in Louisiana makes that species to a worthwhile research object, and even especially being an unwelcome invader, which needs to be better understood to successfully be fought.

As most known ant species, also S. invicta is characterized by hosting a remarkable number of non-ant-inhabitants in their nests, for example mites. One mite species attracted a special attention due to its habit to appear in great numbers on winged ant females and rarely also on their males. Nobody was competent to describe it taxonomically, but I was, that’s why I travelled in 2009 to the Southern Research Station of the USDA Forest Service in Pineville, funded by the German DFG (German Research Foundation). I was very friendly and courteously hosted by the 80 years old colleague John C. Moser, who supported my research by providing access to a microscope and preparation equipment. Although he did not participate directly in my taxonomic work, I honored his contributions of ideas and his interest in my work by offering him a co-authorship.

After the species description was already almost finished and the discussion was about the species‘ name, he unexpectedly insisted in the epitheton „blomquisti“, to honor his assistant Stacy Blomquist. Although he surely was depending on her young dynamic power, I was certainly not amused about this choice. Mrs. Blomquist appeared me being too much involved into the kind of spiritual devoutness, which seemed me to be typical for the whole land. But there was no way out, the species was finally named Histiostoma blomquisti Wirth & Moser, 2010. As a consequence, my name would be forever  connected with hers, an immutable fact. But I still have the freedom to emphasize that I until today think that a biological assistant without a high and internationally well known scientific reputation never deserves a species being named after her. I additionally insist in the fact that I unsolicited would never honor somebody I consider a religious activist with my scientific work. Thus I announce herwith my strict distance to Mrs. Blomquist, who accidentally became namesake of my species.

As all Southern US-States, Louisiana is a land dominated by the creationism, being part of the so called bible belt. I was told that the separation between blacks and whites even goes thus far that there exist black and white churches, but I experienced for sure that Chinese inhabitants are priviledged to be considered white, thus visiting the white churches.

Once I was invited to a private video evening. It was organized by a Chinese assistant of the research institute and a white colleague of him, a hobby marathon runner. They presented the 2003 US/ German/ British co-production „Luther“ with Joseph Fiennes in the main role. Directed by the British Eric Till, this very average movie with a Martin Luther, attractive, slim, completely unlike the historical original, fat with a strong penchant for alcohol, was a strange choice for me as a cineaste, but I expected an entertaining popcorn evening with discussions about good and bad movies. But what I then witnessed was very unexpected.

It was so silent that I could hear the air breath, the flies buzzing around, a mysterious expectation filled the room, while my two hosts stared to the screen, awaiting the first appearance of „Martin Luther“ with religious reverence and deistic adoration. I landed in a private divine service, and when Joseph Fiennes appeared for the first time, both raised their arms ecstatically into the air, praising Luther as the great only one. It kept going on like this, no popcorn for me, but very frequent cigarette breaks instead, I fled outside as often as I could.

It is a well known phenomenon that in areas of unjustice worldwide natives tend to protect their own conscience with an unfounded belief to stand under a special supervision by a god. Only a god of immorality and misanthropy claimes to have created the planet and all life on it within six days, only such a god supports the inequality of races and the discrimination of minorities.

Berlin, November 2017

Histiostoma blomquisti on fire ant queens

Copyrights Stefan F. Wirth

 

Stefan F. Wirth

ich

Stefan Friedrich Wirth is a freelance German biologist, zoologist, evolutionary biologist and acarologist, living in Berlin.

– born in 1972 in the South-West of Germany.

– studies at the FU (Free University) Berlin 1994-2000

– phd thesis at the FU Berlin 2000-2004

– since 2004 research in the fields of systematics, evolution und ecology of mites (Histiostomatidae, Astigmata, Acari)  in cooperation with different international scientific institutions and  videografie/ macro-Videography as documentary contributions, for example to the „arte“-channel documentaries „Voyage sous nos pieds“ by the French director Vincent Amouroux.

selected publications:

Wirth, S. (2003): Das Stammartmuster der Histiostomatidae (Acari) und Beschreibung der durch zwei Männchen-Typen charakterisierten Histiostoma palustre n. sp.. Acarologia 42, 3: 257-270.

Wirth, S. (2004): Phylogeny, biology and character transformations of the Histiostomatidae (Acari, Astigmata). Promotionsarbeit. Internet Publikation, URL:http://www.diss.fu-berlin.de/2004/312.

Wirth, S. (2004): Phylogeny, Morphology and habitats of the Histiostomatidae (Astigmata). Proceedings of the V Symposium of the European Association of Acarologists. Phytophaga, XIV: 389-407.

Wirth, S. (2005): Description of a new species Bonomoia opuntiae (Histiostomatidae, Astigmata) with observations on the function of its eyes. Acarologia, vol. 45, no 4: 303-319. (URL:http://cat.inist.fr/?aModele=afficheN&cpsidt=18276055)

Wirth, S. (2005): Transformations of copulation structures and observations on the male polyphenism in the phylogeny of the Histiostomatidae. Internat. J. Acarol., Vol. 31, No. 2: 91-100.

Wirth, S. (2006): Development of the prelarva and larval behavior to open the eggshell in the Histiostomatidae (Astigmata). Abh. Ber. Naturkundemus. Görlitz 78,1: 93-104.

Wirth, S. (2006): Morphology and function of the gnathosoma in the Histiostomatidae (Astigmata). Acarologia,  vol. 46, no. 1-2: 103-109. (URL:http://cat.inist.fr/?aModele=afficheN&cpsidt=18695493)

Wirth, S. (2007): Phylogeny and characteristic transformations of the Histiostomatidae. In: J.B. Morales-Malacara, V. Behan-Pelletier, E. Ueckermann, T.M. Pérez, E.G. Estrada-Venegas and M. Badii (Eds.), Acarology XI. Proceedings of the XI International Congress of Acarology: 607-615, México.

Wirth, S. & Moser, J.C. (2008): Interactions of histiostomatid mites (Astigmata) and leafcutting ants. In: M. Bertrand, S. Kreiter, K.D. McCoy, A. Migeon, M. Navajas, M.-S. Tixier, L. Vial (Eds.), Integrative Acarology. Proceedings of the 6th Congress of the European Association of Acarologists: 378-384; EURAAC 2008, Montpellier, France.

Wirth S. (2009): Necromenic life style of Histiostoma polypori (Acari, Histiostomatidae). Experimental and applied acarology. DOI number: 10.1007/s10493-009-9295-6. URL:http://www.springerlink.com/openurl.asp?genre=article&id=doi:10.1007/s10493-009-9295-6

Wirth, S. (2010): Food competition and feeding behaviour and its implications for the phylogeny of the Histiostomatidae (Astigmata). – In: Sabelis, M. W. & Bruin, J. (eds.). Trends in Acarology: 37-40.

Wirth S. & Moser J. C. (2010): Histiostoma blomquisti N. SP. (Acari: Histiostomatidae) A phoretic mite of the Red Imported Fire Ant, Solenopsis invicta Buren (Hymenoptera: Formicidae). Acarologia 50(3): 357-371.

DunlopJ. A., Wirth1 S., Penney2 D.,  McNeilA., Bradley3R.S.,  Withers3 P. J.,Preziosi2 R. F. (2011): A tiny phoretic mite deutonymph in Baltic amber recovered by X-ray computed tomography. Biology letters doi:10.1098/rsbl.2011.0923.

Krüger J. & Wirth S. (2011): Life cycle of  Sarraceniopus nipponensis (Histiostomatidae: Astigmata) from the fluid-filled pitchers of Sarracenia alata (Sarraceniaceae). Acarologia 51(2): 259-267.

Koller L., Wirth S. and Raspotnig G. (2012): Geranial-rich oil gland secretions: a common phenomenon in the Histiostomatidae (Acari, Astigmata)? International journal of Acarology 38(5-38): 420-426.

Pernek M.1,2, Wirth S.3, Blomquist S. R.4, Avtzis D. N.5, Moser J. C.4 (2012): New associations of phoretic mites on Pityokteines curvidens (Coleoptera, Curculionidae, Scolytinae). Central European Journal of Biology. Volume 7, Issue 1: 63-68.

Pernek M. (1),  Novak Agbaba S.(1), Lackovic N. (1), Dod(1) N., Lukic I. (2), Wirth S. (3) (2012): The role of biotic factors on pine (Pinus spp.) decline in north dalmatia (croat: uloga biotičkih čimbenika u sušenjuborova (Pinus spp.) na područjusjeverne dalmacije). Šumarski list, 5–6, cxxxvi: 343–354.

Wirth S. (1), Pernek M. (2) (2012): First record of the mite Histiostoma ulmi in silver fir and indication of a possible phoretic dispersal by the longhorn beetle Acanthocinus reticulates. Šumarski list, 11–12, CXXXVI: 597–603.

Wirth S. & Garonna A. P.  (2015): Histiostoma ovalis (Histiostomatidae, Acari) associated with Ips sexdentatus (Scolytinae, Curculionidae, Coleoptera): ecology and mite redescription on the basis of formerly unknown adults and nymphs . International Journal of Acarology DOI: 10.1080/01647954.2015.1050062

WIRTH S., WEIS O., PERNEK M. (2016): A comparison of phoretic mites associated with bark beetles Ips typographus and Ips cembrae from Central Croatia. Šumarski list.

WIRTH S. (2016): Description of developmental instars of Bonomoia sibirica n. sp. (Astigmata: Histiostomatidae) with ecological observations and phylogenetic conclusions. Acarina, December issue.

WIRTH S. F. (2021): Two different forms of cryptic species-complexes in mites of the Histiostomatidae (Astigmata) from bank mud and bark beetle-galleries and their significance for applied biodiversity research. Biologe (ed. Stefan F. Wirth), category : original scientific papers volume 1 (2021; 2022) , 1-7. URL: https://biologe.wordpress.com/2021/12/31/two-different-forms-of-cryptic-species-complexes-in-mites-of-the-histiostomatidae-astigmata-from-bank-mud-and-bark-beetle-galleries-and-their-significance-for-applied-biodiversity-research

WIRTH S. F. (2022): Specific phoretic mites as microclimate originators in special ephemeral soil habitats as presumed co-creators of nutrient-rich soil areas. Poster publication to Global Symposium on Soils for Nutrition, 26/07/2022 – 29/07/2022, by FAO (Food and Agriculture Organization of the United Nations), poster 167. https://www.fao.org/fileadmin/user_upload/GSP/GSOIL4N/GSOIL4N-Posters/ID_167.pdf

Stefan F. Wirth

ich

Stefan Friedrich Wirth ist ein freischaffender deutscher Biologe, Zoologe, Evolutionsbiologe und Acarologe, der jedoch durch Geburt auch über italienische Wurzeln verfügt.  Er lebt in Berlin.

– Geboren 1972 im Südwesten Deutschlands.

– Studium an der FU Berlin 1994-2000

– Promotion an der FU Berlin 2000-2004

– Ab 2004 bis heute: Forschung (bis 2014 auch Lehre an der FU Berlin) in den Bereichen Systematik, Evolution und Ökologie der Milben (am Beispiel bestimmter Milben aus der Gruppe der Astigmata) in Kooperation mit diversen internationalen Forschungseinrichtungen, sowie Videografie/ Makro-Videographie für künstlerisch-dokumentarische Zwecke, z.B. als Beiträge zu den Dokumentarfilmen „Voyage sous nos pieds“ des französischen Regisseurs Vincent Amouroux, die 2016 auf dem französisch-deutschen Fernsehsender „arte“ ausgestrahlt wurden.

Ausgewählte Publikationen:

Wirth, S. (2003): Das Stammartmuster der Histiostomatidae (Acari) und Beschreibung der durch zwei Männchen-Typen charakterisierten Histiostoma palustre n. sp.. Acarologia 42, 3: 257-270.

Wirth, S. (2004): Phylogeny, biology and character transformations of the Histiostomatidae (Acari, Astigmata). Promotionsarbeit. Internet Publikation, URL:http://www.diss.fu-berlin.de/2004/312.

Wirth, S. (2004): Phylogeny, Morphology and habitats of the Histiostomatidae (Astigmata). Proceedings of the V Symposium of the European Association of Acarologists. Phytophaga, XIV: 389-407.

Wirth, S. (2005): Description of a new species Bonomoia opuntiae (Histiostomatidae, Astigmata) with observations on the function of its eyes. Acarologia, vol. 45, no 4: 303-319. (URL:http://cat.inist.fr/?aModele=afficheN&cpsidt=18276055)

Wirth, S. (2005): Transformations of copulation structures and observations on the male polyphenism in the phylogeny of the Histiostomatidae. Internat. J. Acarol., Vol. 31, No. 2: 91-100.

Wirth, S. (2006): Development of the prelarva and larval behavior to open the eggshell in the Histiostomatidae (Astigmata). Abh. Ber. Naturkundemus. Görlitz 78,1: 93-104.

Wirth, S. (2006): Morphology and function of the gnathosoma in the Histiostomatidae (Astigmata). Acarologia,  vol. 46, no. 1-2: 103-109. (URL:http://cat.inist.fr/?aModele=afficheN&cpsidt=18695493)

Wirth, S. (2007): Phylogeny and characteristic transformations of the Histiostomatidae. In: J.B. Morales-Malacara, V. Behan-Pelletier, E. Ueckermann, T.M. Pérez, E.G. Estrada-Venegas and M. Badii (Eds.), Acarology XI. Proceedings of the XI International Congress of Acarology: 607-615, México.

Wirth, S. & Moser, J.C. (2008): Interactions of histiostomatid mites (Astigmata) and leafcutting ants. In: M. Bertrand, S. Kreiter, K.D. McCoy, A. Migeon, M. Navajas, M.-S. Tixier, L. Vial (Eds.), Integrative Acarology. Proceedings of the 6th Congress of the European Association of Acarologists: 378-384; EURAAC 2008, Montpellier, France.

Wirth S. (2009): Necromenic life style of Histiostoma polypori (Acari, Histiostomatidae). Experimental and applied acarology. DOI number: 10.1007/s10493-009-9295-6. URL:http://www.springerlink.com/openurl.asp?genre=article&id=doi:10.1007/s10493-009-9295-6

Wirth, S. (2010): Food competition and feeding behaviour and its implications for the phylogeny of the Histiostomatidae (Astigmata). – In: Sabelis, M. W. & Bruin, J. (eds.). Trends in Acarology: 37-40.

Wirth S. & Moser J. C. (2010): Histiostoma blomquisti N. SP. (Acari: Histiostomatidae) A phoretic mite of the Red Imported Fire Ant, Solenopsis invicta Buren (Hymenoptera: Formicidae). Acarologia 50(3): 357-371.

DunlopJ. A., Wirth1 S., Penney2 D.,  McNeilA., Bradley3R.S.,  Withers3 P. J.,Preziosi2 R. F. (2011): A tiny phoretic mite deutonymph in Baltic amber recovered by X-ray computed tomography. Biology letters doi:10.1098/rsbl.2011.0923.

Krüger J. & Wirth S. (2011): Life cycle of  Sarraceniopus nipponensis (Histiostomatidae: Astigmata) from the fluid-filled pitchers of Sarracenia alata (Sarraceniaceae). Acarologia 51(2): 259-267.

Koller L., Wirth S. and Raspotnig G. (2012): Geranial-rich oil gland secretions: a common phenomenon in the Histiostomatidae (Acari, Astigmata)? International journal of Acarology 38(5-38): 420-426.

Pernek M.1,2, Wirth S.3, Blomquist S. R.4, Avtzis D. N.5, Moser J. C.4 (2012): New associations of phoretic mites on Pityokteines curvidens (Coleoptera, Curculionidae, Scolytinae). Central European Journal of Biology. Volume 7, Issue 1: 63-68.

Pernek M. (1),  Novak Agbaba S.(1), Lackovic N. (1), Dod(1) N., Lukic I. (2), Wirth S. (3) (2012): The role of biotic factors on pine (Pinus spp.) decline in north dalmatia (croat: uloga biotičkih čimbenika u sušenjuborova (Pinus spp.) na područjusjeverne dalmacije). Šumarski list, 5–6, cxxxvi: 343–354.

Wirth S. (1), Pernek M. (2) (2012): First record of the mite Histiostoma ulmi in silver fir and indication of a possible phoretic dispersal by the longhorn beetle Acanthocinus reticulates. Šumarski list, 11–12, CXXXVI: 597–603.

Wirth S. & Garonna A. P.  (2015): Histiostoma ovalis (Histiostomatidae, Acari) associated with Ips sexdentatus (Scolytinae, Curculionidae, Coleoptera): ecology and mite redescription on the basis of formerly unknown adults and nymphs . International Journal of Acarology DOI: 10.1080/01647954.2015.1050062

WIRTH S., WEIS O., PERNEK M. (2016): A comparison of phoretic mites associated with bark beetles Ips typographus and Ips cembrae from Central Croatia. Šumarski list.

WIRTH S. (2016): Description of developmental instars of Bonomoia sibirica n. sp. (Astigmata: Histiostomatidae) with ecological observations and phylogenetic conclusions. Acarina, December issue.

WIRTH S. F. (2021): Two different forms of cryptic species-complexes in mites of the Histiostomatidae (Astigmata) from bank mud and bark beetle-galleries and their significance for applied biodiversity research. Biologe (ed. Stefan F. Wirth), category : original scientific papers volume 1 (2021; 2022) , 1-7. URL: https://biologe.wordpress.com/2021/12/31/two-different-forms-of-cryptic-species-complexes-in-mites-of-the-histiostomatidae-astigmata-from-bank-mud-and-bark-beetle-galleries-and-their-significance-for-applied-biodiversity-research

WIRTH S. F. (2022): Specific phoretic mites as microclimate originators in special ephemeral soil habitats as presumed co-creators of nutrient-rich soil areas. Poster publication to Global Symposium on Soils for Nutrition, 26/07/2022 – 29/07/2022, by FAO (Food and Agriculture Organization of the United Nations), poster 167. https://www.fao.org/fileadmin/user_upload/GSP/GSOIL4N/GSOIL4N-Posters/ID_167.pdf